New large pterosaur tracks from Korea and their implications on terrestrial behavior.

Pterosaurs were important components of Mesozoic ecosystems, occupying diverse ecological niches from the Late Triassic to the end of the Cretaceous. Among them, neoazhdarchians have been hypothesized as terrestrial carnivorous based on anatomical and functional evidence, yet direct indications of predation on land have remained elusive. Here we report Jinjuichnus procerus ichnogen. et ichnosp. nov., a new pterosaur track discovered in the Jinju Formation of South Korea. The track, characterized by a compact and long metatarsus in its pedal configuration, is consistent with neoazhdarchian pterosaurs. The trackway is preserved in close association with a small tetrapod trackway that exhibits abrupt changes in direction and increased stride length. This association provides the potential ichnological evidence of terrestrial vertebrate interaction by a pterosaur. Nonetheless, an alternative interpretation of the two trackways remains possible, making it difficult to confirm any direct interaction between the trackmakers. The paired trackways offer insight into the factors to consider when evaluating potential interactions with the trackmaker. While scenarios such as predation remain ambiguous, they nonetheless highlight the interpretive complexity inherent in assessing behavioral associations preserved in trackway assemblages.