Identifying climate thresholds for dominant natural vegetation types at the global scale using machine learning: Average climate versus extremes.

The global distribution of vegetation is largely determined by climatic conditions and feeds back into the climate system. To predict future vegetation changes in response to climate change, it is crucial to identify and understand key patterns and processes that couple vegetation and climate. Dynamic global vegetation models (DGVMs) have been widely applied to describe the distribution of vegetation types and their future dynamics in response to climate change. As a process-based approach, it partly relies on hard-coded climate thresholds to constrain the distribution of vegetation. What thresholds to implement in DGVMs and how to replace them with more process-based descriptions remain among the major challenges. In this study, we employ machine learning using decision trees to extract large-scale relationships between the global distribution of vegetation and climatic characteristics from remotely sensed vegetation and climate data. We analyse how the dominant vegetation types are linked to climate extremes as compared to seasonally or annually averaged climatic conditions. The results show that climate extremes allow us to describe the distribution and eco-climatological space of the vegetation types more accurately than the averaged climate variables, especially those types which occupy small territories in a relatively homogeneous ecological space. Future predicted vegetation changes using both climate extremes and averaged climate variables are less prominent than that predicted by averaged climate variables and are in better agreement with those of DGVMs, further indicating the importance of climate extremes in determining geographic distributions of different vegetation types. We found that the temperature thresholds for vegetation types (e.g. grass and open shrubland) in cold environments vary with moisture conditions. The coldest daily maximum temperature (extreme cold day) is particularly important for separating many different vegetation types. These findings highlight the need for a more explicit representation of the impacts of climate extremes on vegetation in DGVMs.