A stochastic differential game approach toward animal migration.

A stochastic differential game model for animal migration between two habitats under uncertain environment, a new population dynamics model, is formulated. Its novelty is the use of an impulse control formalism to naturally describe migrations with different timings and magnitudes that the conventional models could not handle. Uncertainty of the environment that the population faces with is formulated in the context of the multiplier robust control. The optimal migration strategy to give the maximized minimal profit is found through a Hamilton-Jacobi-Bellman quasi-variational inequality (HJBQVI). A key message from HJBQVI is that its free boundary determines the optimal migration strategy. Solving the HJBQVI is carried out with a specialized stable and convergent finite difference scheme. This paper theoretically suggests that the sub-additivity of the performance index, the index to be optimized through the migration, critically affects the resulting strategy. The computational results with the established scheme are consistent with the theoretical predictions and support importance of the sub-additivity property. Social interaction to reduce the net mortality rate is also quantified, suggesting a linkage between the present and existing population dynamics models.